What does the X4 mean in an PCIe Gen X4 NVME drive?

What does the X4 mean in an PCIe Gen X4 NVME drive? can you connect pcie 4 0 in 3 0 slot
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can you win twice on the same lottery ticket “PCIe will not be supported on motherboards released prior to the X570, so 400-series and 300-series will have PCIe support,” an AMD

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